Education and Training

               Postdoctorate:                    University of Colorado, Boulder, Colorado, 1980-1984

               Graduate:                            University of Minnesota, St. Paul, Minnesota, 1975-1980

                                                            Ph.D. in Genetics

               Undergraduate:                   Geneva College, Beaver Falls, Pennsylvania, 1971-1975

                                                            B.S. in Biology, summa cum laude

Professional Experience and Appointments

2001-                                Professor, Department of Biology, Haverford College (PA)

2005-2008                       Associate Provost, Haverford College

1998-2001                       Department Chair, Department of Biology, Haverford College (PA)

2013-2014

1995-2001                       Associate Professor, Department of Biology, Haverford College (PA)

1989-1995                       Associate Member, Division of Basic Sciences, Fred Hutchinson Cancer Research Center, Seattle, WA

1985-1989                       Assistant Member, Division of Basic Sciences, Fred Hutchinson Cancer Research Center, Seattle, WA

1986-1995                       Affiliate Faculty, Department of Genetics  University of Washington, Seattle, WA

1980-1984                       Postdoctoral research fellow, Department of Molecular, Cellular, and Developmental Biology, University of Colorado, Boulder,  Supervisor:  W.B. Wood

1980                               Lecturer, Department of Biology, Macalester College, St. Paul, MN

1975-1980                       Predoctoral trainee, Department of Genetics and Cell Biology, University of Minnesota, St. Paul, MN.  Thesis advisor:  R.K. Herman

Selected College-Wide Administrative Service at Haverford College

2012-                              Member, then Chair of Educational Policy Committee

1998-2001                       Department Chair, Department of Biology, Haverford College

2013-2014

2009-2013                       HHMI Program Director 

2009-10                           Chair, search committee for Chief Information Officer, Haverford  College

2007-2008                       Co-chair, Master Planning Committee, Haverford College

2005-2008                       Associate Provost, Haverford College

2004-2005                       Director: Koshland Integrated Natural Sciences Center

2003-2005                       Natural Science Division Representative, Academic Council  

1999- 2003                      HHMI Program Director  (student research, minority outreach, and faculty development)

1997-2002                       Koshland Integrated Science Center Planning and Building Committee-  Biology Department Representative