Robert M. Sapolsky is the senior author on this paper.  When asked what role he played in the study, Dr. Sapolsky responded by emphasizing the indirect role he played as a consultant to Meaney, as opposed to playing a direct role in the way the study was conducted.  According to Sapolsky, Meaney was mainly responsible for running the study.Watch Sapolsky talk about his role in the study. (16:12-16:32)